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This Constitution Class starship under the command of [[Riferimento::Matthew Decker]] served Starfleet for over 20 years, including three five-year exploration missions. Captain Decker was promoted to Commodore in 2263, but remained in command of the [[Riferimento::USS Constellation NCC-1017|Constellation]]. In 2267, the Constellation encountered several star systems where planets had recently been destroyed, and found the cause in System L-374… an ancient alien doomsday machine that was breaking up the planets with an antiproton beam. The ship was badly damaged by the planet killer, and

entire crew killed, except for Commodore Decker. After being rescued, Decker heroically flew a shuttlecraft into the planet killer in an attempt to destroy it.